Copenhagen, Denmark, 1926. Einar Wegener (played by Eddie Redmayne) and his wife Gerda (Alicia Vikander) are a happily married couple. Both are artists, Einar preferring landscapes and she portraits. One day Einar poses for a portrait of Gerda's while wearing a dress. This is initially done as a lark, as is the later attendance at a party dressed as a woman. However, Einar soon discovers that she is in fact a woman and over time prefers being Lili. At first she and Gerda try to have her situation "cured" but this leads nowhere (other than to many doctors trying to have Lili locked up as a pervert and/or lunatic). Her voyage of self-discovery will ultimately lead to her undergoing the first ever sex-change operation.

Theodore is a lonely man in the final stages of his divorce. When he's not working as a letter writer, his down time is spent playing video games and occasionally hanging out with friends. He decides to purchase the new OS1, which is advertised as the world's first artificially intelligent operating system, "It's not just an operating system, it's a consciousness," the ad states. Theodore quickly finds himself drawn in with Samantha, the voice behind his OS1. As they start spending time together they grow closer and closer and eventually find themselves in love. Having fallen in love with his OS, Theodore finds himself dealing with feelings of both great joy and doubt. As an OS, Samantha has powerful intelligence that she uses to help Theodore in ways others hadn't, but how does she help him deal with his inner conflict of being in love with an OS?

Three grown prodigies, all with a unique genius of some kind, and their mother are staying at the family household. Their father, Royal had left them long ago, and comes back to make things right with his family.

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