The film tells the story of Russian émigré, and the only survivor from ship crash Yanko Goorall, and servant Amy Foster in the end of nineteenth century. When Yanko enters a farm, sick and hungry after the shipwreck, everyone is afraid of him, except for Amy, who is very kind and helps him. Soon he becomes like a son for Dr. James Kennedy and romance between Yanko and Amy follows.

On the coast of Cork, Syracuse is a divorced fisherman who has stopped drinking. His precocious daughter Annie has failing kidneys. One day, he finds a nearly-drowned young woman in his net; she calls herself Ondine and wants no one to see her. He puts her up in an isolated cottage that was his mother's. Annie discovers Ondine's presence and believes she is a selkie, a seal that turns human while on land. Syracuse is afraid to hope again.

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